Thursday, 21 August 2014

Taxonomy - Biology Class 11 notes (complete notes upto the point)

KAIZEN CLASSES



                                         PLANT TAXONOMY OR SYSTEMATIC BOTANY


-study of identification, nomenclature and classification of organism.
identification: it is the finding of correct name and place of  an organism in a system of classification
nomenclature: it is the assigning  of proper names to organisms
or it is the determination of correct name as per rules.
classification: it deals with ordering into groups.

-  term taxonomy was given by A.P. de candolle.
v   father of taxonomy & Modern Botany: carolus Linnaeus .
v   Systematics: Taxonomy in which plants are classified on basis of phylogeny (evolutionary history) or it is the study of diversity of organism & all their comparative and evolutionary relationship based on comparative anatomy, comparative physiology, comparative biochemistry.


SYSTEMATICS = TAXONOMY +PHYLOGENY + DIVERSITY
Term “Systematic ’’ was given by Linnaeus
 Term “Phylogeny’’ was given by Lamarck

                       DIFERENT BRANCHES OF FAXONOMY
1.        ALPHA TAXONOMY:
v           Early phase of taxonomy /Descriptive Systematics
v           In this taxonomy plants are classified on the basis of only
              morphological characters
2.        BETA TAXONOMY: It deals with the arrangement of taxa into
                                       hierarchial system.
3.       BIOSYSTEMATICS / NEO – SYSTEMATICS / NEW SYSTEMATICS /
                                       OMEGA  TAXONOMY:
v           It involve not only morphological character but also genctical,
             anatomical, physiological, palynological character i.e. in this
             Systematics all biological parameters are taken into consideration to 
             classify organism.
v           Term Neo- Systematic was given by Julian Huxley Q.

             It led to the origin of different branches-
a.        Cytotaxonomy / Karyotaxonomy – Taxonomy + Cytology
v           plant are classified on basis of cytological characters like number,
             shape, size of chromosomes and their behavior during cell division                                 
             etc. etc.
     b.  Chemo Taxonomy / Bio chemical Taxonomy:
v   It deals with classification of plant on basis of chemical characters like  latex, gum, resin, oil
c. Numerical / phonetic /Adansonian Classification:
v   Proposed by Adanson
- It employs numerical methods for the evaluation of similarities and
         differences between groups of organism.
         -In this method as many characteristics as possible are used for these
         Comparisons , without extra emphasis being given to any one.
         -  All characters considered for analysis are given equal importance and
         weightage.
-These data are processed and analysed though computers.
         -This system is considered better because it uses large number of 
         comparable characters to assign a place to the species.
v   Sokal and sneath wrote the book “principles of Numerical taxonomy’
  d. Experimental Taxonomy:
         It is based on experimental determination of genetic inter relationship  identification of evolutionary units (subspecies / varieties) within a species.
 4.   TAXONOMY:
v              Based upon phylogeny 
v              Term ----- taxonomy were given by Turill.
 5. CLASSICAL / OLD SYSTEMATICS:
v  Its concept was given by Plato. Aristotle & followed by Linnaeus.
v  According to this concept species is static unchangeable. so instead of large population a few
     Individual can be studied to get information about all characters of species.
v  This is called typological consent of species.


6. CLASSICAL TAXONOMY:
v       Based on origin, evolution and variation of organism.

                                          History of classification:
Chandyogya Upanishad classified animals into three categories viz.
                       jivaja (viviparous), Andaja (oviparous) and Udbhija (vegetal origin).
susuruta samhita (600 B.C.,written by susuruta – father of surgery) classified all substances into
                             Sthavara (immobile) eg: plants
                             Jangama (mobile) e.g. animals
parasra grouped angiosperms into –
                            Dvimatruka (Dicots)
                            Ekamatruka (Monocots)   
Hippocrates (father of Medicine) and Aristotle arranged animals into 4major groups like- Insects, Birds, Fishes & Whales.
Aristotle wrote Historia Animals (the first book of zoology). He arranged animals into Hierarchies. He formulated the Great Chain of Being   (Scala Naturae i.e. Ladder of Nature)- a chain of progressive Changes in nature(some type of evolution).
John Ray  described 18,000 plants and animals in Historia Generalis Plantarum.
 He coined the term genus and species.

                              CONCEPT OF SPECIES

Species: Group of individuals, Which can interbreed & produces fertile offspring.
v       Term species was given by John Ray.
Reproductive Isolation: Members of diff species can’t interbreed naturally. This is called reproductive isolation. But they can interbreed                                     in captivity.

 Male   Horse   X   Female donkey    Female Horse      X      male donkey
(Equus equus )     ( Equus asinus)

                         Mule                                                  Hinny

v  Reproductive isolation maintains sanctity or purity of species.

Biological Concept of species: Biological concept of species was given by Adolf Mayr.It is based on reproductive isolation.
Nominalistic Concept of Species: According to this concept Sps. have no existence only individuals exist.
Morphospecies:                  Morphologically similar species.
Taxonomic Species:           Species have definite binominal name. 
Sibling Species:                  Morphologically similar and reproductively isolated species
Allopatric species:              species living in different geographical area are called allopitic
                                              species.
Sympatric Species:            Species living in same geographical area are called Allopatric
                                              species .  
Neontological species:       living Species.
Palaeospecies :                   Fossil species.
Monotypic Species:           Species having one or no variety or subspecies.
Polytypic Species:             Species having many varieties or subspecies
Monotypic Genus:            Genus having only one species
                                            e.g. Fritschiella tuberosa (terrestaial alga), Welwitschia ,  mirabilis.

TAXONOMIC  HIERARCHY
Species – Is the smallest unit of classification.
Genus (pl. Genera) – Group of similar species Term genus was given by JOHN RAY
eg: potato & Brinjal constitue genus Solanum
                                                                                                            
Family – Group of closely related genus.
                         eg: Solanum, Lycopersicon, capsicum are included in family
                         Solanaceae
Order – Group of similar families.
Class   -  Group of similar order.
Division / phylum (pl. phyla) – Group of similar classes.
kingdom – Group of similar phyla.
      These categories are ranked above the other. This is called Hierarchical
      classification.
     The higher the categories lesser will be the no. of common characters tics of
     organism
v      Kingdom is largest unit of classification.
v      There are 7 obligate categories.
v      Each category can be divided into sub category.
v      phylum of animal is equivalent to division in plant.
v      Term phylum was given by cuvier.
Taxon:    It is the taxonomic group of any rank.
               Term taxon was given by Adolf Mayr for animals by J.Lam for plants.
Suffixes used in different taxon. 
ICBN-    According to ICBN (international code of Botanical nomenclature) the names of diff. taxon must end in following suffixes.
               Rank of category                                   Suffixes    
  1.  Phylum / division                                    1.-phyta (eg. Anthophyta)
  2.  sub phylum / sub div.                              2.-phytina
  3. class                                                         3.-opside (Magaoliopsica)
  4. Sub class                                                  4.-ideae
  5. Order                                                       5.-ales (eg. Malvales)
  6. Sub-order                                                 6.-ineac
  7. family                                                       7.-aceae
  8. sub-family                                                8.-oideae
  9. Tribe                                                         9.-eae
10. Sub-tribe                                                10.-inae

                                Name of 8 families
 Old Name                                                        New Name
  1. Cruciferae                                                      1.Brassicaceac
  2. Compositae                                                    2.Asteraccae
  3. Graminae                                                       3.poaccae
  4. Labiatae                                                         4.Lamiaceae
  5. Umbelliferae                                                   5.Apiaceae
  6. Leguminoseae                                                6.Fabaceae
  7. palmae                                                            7.Arccaccae
  8. Gutifereae                                                       8. Clusiaccae
The rules of nomenclature are framed and standardised by ICBN & ICZN
Foundation of ICBN appeared in 1978
ICZN- International code of Zoological nomenclature 
ICVN – International code of Viral nomenclature
ICNCP- International code of Nomenclature for Cultivated Plants

                              SYSTEMS OF NOMENCLATURE
1. polynomial Nomenclature:
                        According to this system name of organism consist of many  Words.                                                                                                                                  
     It was cumbersome so was rejected later
                         eg: Caryophyllum saxatilis foils gramineus umbellatus corymbis
2. Bionomial Nomenclature :
                       -The idea of binomial nomenclature was given by Gaspard Bauhin in his  book  PENAX.
                        -But this was elaborated and employed by Carolus Linnaeus in his book “species plantarum’’.  
                         -Acc. to this system name consist of two Latin words first word is generic epithet & 2nd is species or specific epithet .
                         -Generic name is a noun so started with capital letter and specific name is an adjective so is Written is small letters.
                         -Both generic & specific name are Written in italics or separately underlined to indicate their Latin origin.
                          - If specific name consist of two words then they are hyphened.
                              eg. Hibiscus rosa – sinensis.
                          -Biological name are derived from Latin language or Latinized.
                         -Latin was the common language among the scholars of that time. This language is still recommended in nomenclature because now it is a dead language and it is not changing or evolving in form or spelling.
             Authority of the name :
                         -The abbreviated name of scientist written after botanical name is called the authority of the name . It is the name of scientist who proposed name & described the species. It is written in Roman & not underlined
                             eg. Rosa indica linn .
            Parenthetical Authority :
                         -When some changes are made in botanical name, then the name of original author is placed in parenthesis (bracket) and is followed by name of person, who made changes.
                           eg. Albizza lebbek (Linn.) Benth.
          Principle of priority :    
                       -According to this rule each taxon can bear one correct name and correct name is earliest legitimate one.
        Tautonyms:
                       -When specifies name exactly repeats generic name
                            eg. Naja naja
                               Bubo bubo (owl)
                               Eqqus equus (horse)
                        -These are not applicable (illegitimate) in botanical nomenclature, but applicable in zoological nomenclature.                   
 3. Trinomial Nomenclature:
                         -According to this system name of organism consist of three words. First word is genus, second is species and third is variety or subspecies.
                               eg: Brassica oleracea var. botrytis
                                     Homo sapiens sapiens
        Typification : 
                         The name of taxon are determined by means of nomenclature type / type genus.
        Nomenclatural type / type genus:  is that element with which the name of taxon is permanently attached.
                   Nomenclatural types are of following type
      Holotype :    it is the  original specimen which is designated as nomenclatural type.
      Isotype :       Duplicate of Holotype
                            if several plants are collected at the same time then one is Holotype and other are Isotype.                                                                   
  
Lectoype: When Holotype is missing then specimen selected from original  material (isotypes) to serve as nomenclatural type is called lectotype.
Neotype: When all the original material on which the description of species  Was based is missing then new specimen selected to serve as  nomenclatural type is called Neotype.
                             
                   SYSTEMS OF CLASSIFICATION
v     placing of an organism into a proper category is called classification
v     Organisms are classified to establish relationship.
         There are 3 systems of classification
1. Artificial system :
        In this system plants are classified on basis of a few (on or two) morphological characters.
a.       Theophrasturs (300BC) He 1st of all classified the plants into four categories Viz-herbs shrubs, Undershrubs &                                  trees in his book “Historia plantarum’’
b.       Carolus Linnaeus: He was born on 23 may 1707 at Result village, South Sweden.
                      Books of Linnaeus:
             1.Species Plantarum          2.Genera plantarum
3. Classes plantarum        4. Philosophia Botanica
5. Critiea plantarum           6. Hortus Upplandicus
7. Hortus Cliffortinus         8. Systemea Naturae in1758AD. for animal classification)
      -Linnaeus classified all the plants into 24 classes on basis of sex orgar s. So his classification is also called sexual system of classification
                         24    classes
       1.  Monandria – 1 stamen                   2.   Diandria – 2 stamen
       3.  Triandria – 3 stamen                     4.   Tetrandria – 4 stamen
       5.  Pentandria – 5 stamen                  6.   Hexandria – 6 stamen
       7.  Heptandria – 7 stamen                  8.   Octandria – 8 starmen
       9.  Enneandria  - 9 stamen                 10. Decandria – 10 starmen
 11. Dodecandria – 11 to 19 stamen free
 12. Icosandria – 20 or more stamens but attached to calyx
 13. polyandria- 20 or more stamen but free
 14. Didynamia – Didynamous flowers
 15. Tetradynamia – tetradynamous
 16. Monoadelphia – monadelphous
 17. Diadelphia diadelphous
 18. Polyadelphia polyadelphous
 19. Syngenesia – Syngenesuous
 20. Gynandria – Cryandeous
 21. Monoecia – Monocecious
 22. Dioecia
 23. Polygonia
 24. Crytogamia
                                                      Outline of Classification
                                                                 Plant kingdom
                
                          
                     Cryptogams                                                        Phanerogams or Spermatophytes
                                                                                                     (Seed beaing plants)
  
                         
                          Dicotyledonae               Gymnosperms                             Monocotyledonae
 
         
      Polypetalae                    Gamopetalae                          Monochlamydoe/Incomplete
       (Petals free)                     (Petals fused)                                  (One whorl of perianth)
                                
                              Inferae        Heteromerae       Bicarpillate
                                                                                                                    
                                                                                                          Divide into 8 series
                                                                                                               
Thalamiflorae               Disciflorae                      Calyciflorae
  (Hypogynous)        A disc (cushion shaped        (epigynous or              Families
                                 Is present at the base of          Perigynous)                                  Divided into
                                  Ovary, Hypogynous)                                                                  7 – series


 
divided into
6 orders                           4 Orders                        5 Orders
eg: Ranunclaecae             eg Rutaceae                 eg Leguminoseae
Cruciferae, Papaveraceae                                              Cucurbitaceae
        Malvaceae   

2. Natural System of Classification
     In this plants are classified on basis of all available morphological      characters
    eg: Bentham & Hooker’ s classification

          Bentham & Hooker were British botanist associated with Royal Botanical Garden, Kew . They proposed natural  system of classification in their book “Genera plantarum’’(3 volumes) in this classification they classified 7569 gnera & 97205 species in 200 families. This classification is post Darwinian but it is not influenced by evolution.


v   This System in followed is Britain and its Commonwealth countries like India,
       But is not followed in America & Germany, they follow Engler and prantl’s classification.

Draw Backs of Bentham & Hooker Classification
1.      The placed (sand witched) gymnosperm b/w Dicots and monocots
2.      Family cucurbitaceae (order passiflorales) is having gamcpetlous  condition but has been placed in Polypetalae.
3. Phylogenetic  Classification / Chadisties / phyletic :
v      Plants are classified on the basis of evolutionary history.
        Supporters : Engler & Prantl, Hutchinson, Bessey, Ostwald – Tipo, Takhtajan, Cronquist etc.
             
              Engler & Prantl’s Classification (German Botanist)
·         They published their classification in  Die Naturlichen pflenzen familien’’.
·         They classified all the plants in 280 families.



                                                              Plant Kingdom
                                            
   Nom Embroyphyta                                                                                Embryophyta              (no embryo formation)
     eg. Algae, fungi                                    
                                       Embryophyta Asiphonogama                  Embryophyta siphonogama
                                       Pollen tube absent                                           - Pollen tube present
                                      eg. Bryophyta & Pteridophyta
 
                                                                        Gymnosperm                                  Angiosperm
                                                                         

                                                                                  Monocots                                  Dicots


 
                                                                                   11 – Orders


 


                                                 Arehichlamydae                            Metachlamydae / Sympetalae
                                                  eg. Apetalous or                                 eg. Gamopetalous
                                                       Polypetalous     
Draw Backs
     1.  They placed monocots before Dicots.
     2.  They classification is partly phylogentie & partly natural. 

         Hutchinson’s Classification
-        British Botanist
-        Published classification in his book “The families of flowering plants’’
-        There are 110 order & 411 families in his classification
                                                        Phanerogams
 
        Angiosperms                                                                            Gymnosperms 


 


       Dicots                                                    Monocots


 


                              Lignosae (woody plans)                     Calyciferae – calyx & corolla distinct
                       
                              Herbaceae                                          Corolliferae – calyx & corolla more or
                                                                                                              less similar

                                                                                        Glumiflorae – Perianth reduced

                  Takhtajan: used the term magnoliophyta for Angiosperms.


Some primitive and advanced characters
           (Acc. to I Iutchinson & Bessey)
            Primitive                                                                 Advanced
1.      Trees & Shrubs                                                        1.Herbal
2.      Perrennial & biennial                                              2. Annuals
3.      Alternate phyllotaxy                                                3. Opposite Phyllotaxy
4.      Simple leaf                                                               4. Compound leaf
5.      Bisexual flower                                                       5. Unisexual
6.      Solitary flower                                                        6. Inflorescence
7.      More petals, sepals stamens, etc & free                 7. Less petal, etc & fused
8.      Free carpel (apocarpous)                                        8. Fused carpel’s (syncarpous)
9.      Simple fruit                                                             9. Composite & Aggregate                                                                 
10. Hypogyny                                                              10. Perigyny & Epigyny
11.Endospermic                                                          11. Non endospermic seeds
12. Cotyledons two                                                     12. One
13. Integuments two                                                    13. Integument one
14. Chromosome number low                                    14. Chromosome number high

v       Largest & Highly evolved / advanced family – compositae (Dicots)
v       Most primitive family Ranunculaeeae  (herbaceous gp) and Magnoliaceae (woody gp)
Taxonomic Aids:
  (A) Herbarium : Iy is a Store house of collected plant specimen that are dried, pressed
                              plants  and preserved on sheets. These sheets are arranged according to an
                              accepted system of classification size of herbarium sheet
                              = 28 x 42 cm / 16 ½ ‘’ x 11½ ’’
(B) Botanical Gardens : Place where living plants are grown for reference
                                           - Also called Living Herbaria .
          special sections of Botanical Gardens :
          Caetorium : part of garden where main collections are cacti.
          Orctorium : part of garden where main collection are orchids.
          Bambusetum : part of garden where main collection are bamboos.
          Arboretum : part of garden where main collection are woody plant species (trees&
                                  shrubs)
         Pinetum :  It is an arboretum where main collection are conifers
         Palm-house : palms are grown.
         Largest herbaria & botanical garden of would : Royal Botanical Garden, Kew,
                                                                        England. also called Botanical capital of world
          Started by sir Henry Capel, then development by princess Augusta.        
  Largest herbarium & botanical garden of India – Indian Botanical Garden, Sibpur,
                                                                            Hawrah Kolkata.
         It was started by Robert Kyd, organized by William Roxburghii .
NBRI – National Botanical Research Institute,  Lucknow.
                    Built by Nawab saadat Ali khan (1789-1814) & adapted by Wajid Ali Shah.
FRI – Forest Research Institute Dehradun.
Lal Bagh Botanical Garden- Banglore.
Lloyd Botanical Garden- Darjeeling
Botanical Garden, Saharanpur
(C) Museum: they have collection of preserved plant and animal specimens for study and
 reference. Specimens are preserved in containers or jars in preservatives (like formalin).
 Larger animal like birds and mammals usually stuffed and preserved.
(D) Zoological parks or Zoos : These the places where wild animals are kept in protected environment under human management. It enable us to learn about their habits and behavior.
(E) Monograph : It is a book / essay which gives comprehensive account of all the available information about a taxon (genus/family etc.) It is highly useful taxonomic aid.
(F) Manual : It is a handbook providing compiled information about a particular with all the families genera, spa and keys for their identification .
(G) Keys : These are artificial analytical  devices in the form of dichotomous table of alternate characteristics which are used in the identification of a class, genus and spa.
                   Keys are two types:
        (i)   Indented or Yoked Key : Here there is a sequence of two or mere  alternate characteristics from which selection or choice is to be made though selection and climination.
       (ii)   Bracketed Key : The key does not separate contrasting characters by subdividing but instead give them numbers in bracket.

      Some Famous Research Institute
1.       CSIR  - Council of scientific and industrial Research,  New Delhi
2.       CDRI – Central Drug& Research Institute, Lucknow
3.       ITRC –  Industrial Toxicological Research, Lucknow
4.       CIMAP – Central Institute, of Medicinal and Aromatic Plant, Lucknow
5.       BSRIP – Birbal Sahni Research Institute of paleo Botany , Lucknow
6.       CTRI – Central Tobacco Research Institute, Rajmundhary, Andhra Pradesh  
7.       CFRI – Central Fruit Research Institute, Sobour, Bhagalpur, Bihar
8.      CFTRI – Central Food Technological Research Institute, Mysore, Karnataka
9.      SBRI –Sugarecame Breeding Research Institute, Coimbatore, T.N.
10.  CRRI – Central Rice Research Institute, Cuttack, Orissa
11.  IRRI – International Rice Research Institute Manilla, Philippines.
12.  ICAR– Indian Council of Agricultural Research Pusa, New Delhi.
13.  ICRI – Indian Agricultural Research Institute, Pusa, Delhi.
14.  NDRI – National Dairy Research Institute, Karnal, Haryana
15.  IVRI – Indian Veterinary Research Institute, Izzatnagar, Barielly
16.  IGFRI – Indian Grassland and Fodder Research Institute, Jhansi
17.  ICRISAT – International Crop Research Institute for semi Arid Tropics, Hyderabad
18.  CAZRI – Central Arid Zone Research Institute, Jodhpur, Rajasthan
19.  CPRI – Central Potato Research Institute, Shimla
20.  CSSRI – Central Soil Salinity Research Institute, Karnal
21.  FAO –  Food and Agricultural Organization.
22.  BSI – Botanical Survey of India, Head office at, Howrah.

Floral Formula : Short hand representation of important floral charactersties
   Br -   Bracteate                                       K – Calyx, 5 sepals, free (polysepalous) 
  EBr – Ebraeteate                                     K – Sepals, tused 
  Brl – Bracteolate                                    (  ) - Fused
 Ebrl – Ebractcolate                                  C – Corolla, 4 petals, free (polypetalous)
            Actinomorphic                              A – Androecium, 10 stamens 
            Zygomorphic                                 C A  Epipetalous
            Bisesual                                         G    Gynaceium , 2 ear pels (free)
            Male                                               G    Superior ovary, hypogynous flower
            Female                                           G    Inferior ovary, Fpigynous flower
            Epicalyx                                         G    Perigynous, semi – Superior semi – inferior.